[文法] 請教這一句的文法及語意

[kfceric (酷內)]

Missing until recently were fossils clearly intermediate, or transitional,
between land mammals and cetaceans.

想請教各位大大
這句話的完整語意為何
以及是否又用到倒裝句的文法
煩請解惑
感激不盡~~

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